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Se realiza gracias a los llamados fotosistemas, que se encuentran en la membrana de los tilacoides en los cloroplastos. Hay dos tipos de fotosistemas: - Fotosistema I , que se encuentra sobre todo en los tilacoides de estroma, y cuyo pigmento diana es la clorofila P El proceso de la fase luminosa, supuesto para dos electrones, es el siguiente: Los fotones inciden sobre el fotosistema II, excitando y liberando dos electrones, que pasan al primer aceptor de electrones, la feofitina.
Los electrones y los protones pasan al complejo de citocromos bf, que bombea los protones al interior del tilacoide. Los electrones de los citocromos pasan a la plastocianina, que los cede a su vez al fotosistema I.
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